Gotta merge :(

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j-hartling
2026-02-05 09:58:56 +01:00
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@@ -193,6 +193,17 @@
year={2002}, year={2002},
}# Cited }# Cited
@article{gray1960fine,
title={The fine structure of the insect ear},
author={Gray, Edward George},
journal={Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences},
volume={243},
number={700},
pages={75--94},
year={1960},
publisher={The Royal Society London}
}# Cited
@article{greenfield1993acoustic, @article{greenfield1993acoustic,
title={Acoustic dueling in tarbush grasshoppers: settlement of territorial contests via alternation of reliable signals}, title={Acoustic dueling in tarbush grasshoppers: settlement of territorial contests via alternation of reliable signals},
author={Greenfield, Michael D and Minckley, Robert L}, author={Greenfield, Michael D and Minckley, Robert L},

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\endentry \endentry
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<bcf:citekey order="69" intorder="1">fisch2012channel</bcf:citekey> <bcf:citekey order="69" intorder="1">clemens2010intensity</bcf:citekey>
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[313] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 157, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 157, warning: 6 characters of junk seen at toplevel
[313] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 178, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 178, warning: 6 characters of junk seen at toplevel
[313] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 196, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 196, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 207, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 207, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 218, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 218, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 227, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 229, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 236, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 238, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 247, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 247, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 256, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 258, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 267, warning: 6 characters of junk seen at toplevel [307] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 267, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 278, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 278, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 287, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 289, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 306, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 298, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 315, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 317, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 378, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 326, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 405, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 389, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 414, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 416, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 433, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 425, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 460, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 444, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 469, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 471, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 504, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 480, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 513, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 515, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 534, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 524, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 543, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 545, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 554, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 554, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 565, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 565, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 574, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 576, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 597, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 585, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 626, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 608, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 636, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 637, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 645, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 647, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 666, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 656, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 687, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 677, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 696, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 698, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 705, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 707, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 722, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 716, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 731, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 733, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 756, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 742, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_B9EJ/347c261ec4135a5723bef5c751f5078f_26287.utf8, line 773, warning: 6 characters of junk seen at toplevel [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 767, warning: 6 characters of junk seen at toplevel
[314] Biber.pm:133> INFO - WARNINGS: 52 [308] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_MFQC/347c261ec4135a5723bef5c751f5078f_8672.utf8, line 784, warning: 6 characters of junk seen at toplevel
[308] Biber.pm:133> INFO - WARNINGS: 53

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@@ -1,14 +1,14 @@
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@@ -131,10 +131,10 @@
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"figures/fig_auditory_pathway.pdf_tex" 1769682029.31397 2624 f8d79af301f26c4639670ba4283887a2 "" "figures/fig_auditory_pathway.pdf_tex" 1769682029.31397 2624 f8d79af301f26c4639670ba4283887a2 ""
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This is pdfTeX, Version 3.141592653-2.6-1.40.25 (TeX Live 2023/Debian) (preloaded format=pdflatex 2025.10.28) 2 FEB 2026 17:12 This is pdfTeX, Version 3.141592653-2.6-1.40.25 (TeX Live 2023/Debian) (preloaded format=pdflatex 2025.10.28) 5 FEB 2026 09:58
entering extended mode entering extended mode
restricted \write18 enabled. restricted \write18 enabled.
file:line:error style messages enabled. file:line:error style messages enabled.
@@ -677,28 +677,33 @@ Overfull \hbox (50.3275pt too wide) in paragraph at lines 192--253
\OT1/cmr/m/n/12 ets ([]; []) and grasshop-pers ([]; \OT1/cmr/m/n/12 ets ([]; []) and grasshop-pers ([];
[] []
[3] [4] [3]
Overfull \hbox (13.04964pt too wide) in paragraph at lines 320--322 Overfull \hbox (149.87093pt too wide) in paragraph at lines 285--304
\OT1/cmr/m/n/12 the neu-ronal sub-strate of song recog-ni-tion ([]; [];
[]
[4]
Overfull \hbox (13.04964pt too wide) in paragraph at lines 318--320
[]\OT1/cmr/m/n/12 Similar re-sponse/filter prop-er-ties within re-cep-tor/interneuron pop-u-la-tions ([]Clemens, Kutzki, []\OT1/cmr/m/n/12 Similar re-sponse/filter prop-er-ties within re-cep-tor/interneuron pop-u-la-tions ([]Clemens, Kutzki,
[] []
[5] (./figures/fig_auditory_pathway.pdf_tex (./figures/fig_auditory_pathway.pdf_tex
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File: figures/fig_auditory_pathway.pdf Graphic file (type pdf) File: figures/fig_auditory_pathway.pdf Graphic file (type pdf)
<use figures/fig_auditory_pathway.pdf, page 1> <use figures/fig_auditory_pathway.pdf, page 1>
Package pdftex.def Info: figures/fig_auditory_pathway.pdf , page1 used on input line 56. Package pdftex.def Info: figures/fig_auditory_pathway.pdf , page1 used on input line 56.
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LaTeX Warning: Text page 6 contains only floats.
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@@ -282,34 +282,32 @@ within the auditory pathway.
\section{Developing a functional model of the\\grasshopper song recognition pathway} \section{Developing a functional model of the\\grasshopper song recognition pathway}
The essence of constructing a functional model of a sensory processing system The essence of constructing a functional model of a given system is to gain a
is to gain a sufficient understanding of the system's essential structural sufficient understanding of the system's essential structural components and
components and the functional roles they might fulfill; and to then build a their presumed functional roles; and to then build a formal framework of
formal framework of manageable complexity around these two aspects. manageable complexity around these two aspects. Anatomically, the organization
Anatomically, the organization of the grasshopper song recognition pathway can of the grasshopper song recognition pathway can be outlined as a hierarchical
be outlined as a hierarchical feed-forward network of three consecutive feed-forward network of three consecutive neuronal
neuronal populations~(Fig.\,\ref{fig:pathway}a-c): Peripheral auditory receptor populations~(Fig.\,\ref{fig:pathway}a-c): Peripheral auditory receptor neurons,
neurons, whose axons enter the ventral nerve cord at the level of the whose axons enter the ventral nerve cord at the level of the metathoracic
metathoracic ganglion; local interneurons that remain exclusively within the ganglion; local interneurons that remain exclusively within the thoracic region
thoracic region of the ventral nerve cord; and ascending neurons projecting of the ventral nerve cord; and ascending neurons projecting from the thoracic
from the thoracic region towards the supraesophageal region towards the supraesophageal ganglion~(\bcite{rehbein1974structure};
ganglion~(\bcite{rehbein1974structure}; \bcite{rehbein1976auditory}; \bcite{rehbein1976auditory}; \bcite{eichendorf1980projections}). The input to
\bcite{eichendorf1980projections}). The input to the network originates from the network originates at the membrane of the tympanal organ, which acts as the
primary sound receiver and is coupled to the dendritic endings of the receptor
neurons~(\bcite{gray1960fine}). The outputs of the network converge somewhere
in the supraesophageal ganglion, which is presumed to harbor the neuronal
substrate of song recognition~(\bcite{romer1985responses};
\bcite{ronacher1986routes}; \bcite{bauer1987separate};
\bcite{bhavsar2017brain}).
Functionally, the ascending neuron population is characterized by a marked
The input to the network originates from increase in response heterogenity compared to the preceding receptor neurons
sound-induced vibrations of the tympanal membrane on each side of the thorax, and local interneurons, which exhibit relatively homogeneous response
which are transduced into electro-chemical signals by the receptor neurons. The properties across their respective populations~(\bcite{clemens2011efficient}).
output from the network converges somewhere in the supraesophageal ganglion, Based on these considerations, the organisation of the model
where the recognition of conspecific songs is presumed to take
place~(\bcite{romer1985responses}; \bcite{ronacher1986routes};
\bcite{bauer1987separate}; \bcite{bhavsar2017brain}). Functionally, the
ascending neuron population is characterized by a marked increase in response
heterogenity compared to the preceding receptor neurons and local interneurons,
which exhibit relatively homogeneous response properties across their
respective populations~(\bcite{clemens2011efficient}). Based on these
considerations, the organisation of the model
pathway~(Fig.\,\ref{fig:pathway}d) comprises two distinct overall stages: pathway~(Fig.\,\ref{fig:pathway}d) comprises two distinct overall stages: