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@@ -81,60 +81,61 @@ distributed accumulation of anatomical, physiological and ethological evidence.
 This process is undoubtedly without alternative; however, it leaves us with the
 challenge of integrating the available fragments into a coherent whole in order
 to address issues such as the interaction between individual system components,
-the functional limitations of the system overall, or taxonomic comparisons of
-systems that process the same sensory modality. Any unified framework that
-captures the essential functional aspects of a given sensory system thus has
-the potential to deepen our current understanding and fasciliate systematic
+the functional limitations of the system overall, or taxonomic comparisons
+between systems that process the same sensory modality. Any unified framework
+that captures the essential functional aspects of a given sensory system thus
+has the potential to deepen our current understanding and fasciliate systematic
 investigations. However, building such a framework is a challenging task. It
 requires a wealth of existing knowledge of the system and the signals it
 operates on, a clearly defined scope, and careful reduction, abstraction, and
-formalization of the underlying anatomical structures and physiological
-mechanisms.
+formalization of the underlying structures and mechanisms.
 
 One sensory system about which extensive information has been gathered over the
 years is the auditory system of grasshoppers~(\textit{Acrididae}). Grasshoppers
-rely on auditory processing primarily for intraspecific communication, which
+rely on their sense of hearing primarily for intraspecific communication, which
 includes mate attraction and evaluation~(\bcite{helversen1972gesang}), sender
 localization~(\bcite{helversen1988interaural}), courtship display~(SOURCE),
 rival deterrence~(\bcite{greenfield1993acoustic}), and loss-of-signal predator
-alarm~(SOURCE). The different behavioral contexts are met with 
+alarm~(SOURCE). In accordance with this rich behavioral repertoire,
+grasshoppers have evolved a variety of sound production mechanisms to generate
+acoustic communication signals for different contexts and ranges using their
+wings, hindlegs, or mandibles~(\bcite{otte1970comparative}). Among the most
+conspicuous acoustic signals of grasshoppers are the species-specific calling
+songs, which broadcast the presence of the singing individual --- usually the
+males of a species --- to potential mates within range. These songs are usually
+more characteristic of a species than morphological traits.
 
-Different acustic signals are used for different behavioral
-contexts and communication ranges
+This reliance on species-specific acoustic signals for mediating
+reproduction represents a strong evolutionary driving force, that resulted in a
+massive species diversification~(\bcite{vedenina2011speciation},
+\bcite{sevastianov2023evolution}). 
 
-Depending on the behavioral context and the communication range,
+The characteristic calling songs are
+produced by stridulation, during which the grasshopper pulls the serrated
+stridulatory file on its hindlegs across a resonating vein on the
+forewings~(\bcite{helversen1977stridulatory},
+\bcite{helversen1997recognition}).
 
-
-Grasshoppers generate their most conspicious acoustic signals
----~commonly referred to as "songs"~--- by stridulation.
-
-
-Different acoustic signals may be generated using different
-body parts ---~wings, hindlegs, or mandibles~--- 
-
-
-Different acoustic signals may be generated using different
-body parts ---~wings, hindlegs, or mandibles~--- but the most conspicious
-
-
-The required acoustic signals for different contexts and ranges
-may be generated using different body parts ---~wings, hindlegs, or
-mandibles~--- but the most common sound production mechanism is stridulation,
-during which the animal pulls the serrated stridulatory file on its hindlegs
-across a resonating vein on the forewings. The resulting "songs"
-
-
-The reliance on acoustic communication signals represents a strong evolutionary
-driving force, that resulted in a massive species diversification among
-grasshoppers~(\bcite{vedenina2011speciation},
+Such elaborate acoustic behaviors co-depend not only on
+reliable auditory perception but also on suitable acoustic signals for
+different contexts and ranges. To this end, grasshoppers have evolved a variety
+of sound production mechanisms using their wings, hindlegs, or
+mandibles~(\bcite{otte1970comparative}). The most conspicuous acoustic signals
+--- the characteristic calling songs --- are produced by stridulation, during
+which the grasshopper pulls the serrated stridulatory file on its hindlegs
+across a resonating vein on the forewings~(\bcite{helversen1977stridulatory},
+\bcite{helversen1997recognition}). The reliance on species-specific acoustic
+communication signals represents a strong evolutionary driving force, that
+resulted in a massive species diversification~(\bcite{vedenina2011speciation},
 \bcite{sevastianov2023evolution}).
 
 
-
-Grasshoppers produce their most conspicious acoustic signals
----~commonly referred to as "songs"~--- by stridulation, during which the
-animal rubs the serrated stridulatory file on its hindleg across a resonating
-vein on the forewing. 
+The most conspicuous acoustic signals
+--- the characteristic calling songs --- are produced by stridulation, during
+which the grasshopper pulls the serrated stridulatory file on its hindlegs
+across a resonating vein on the forewings~(\bcite{helversen1977stridulatory},
+\bcite{helversen1997recognition}). Grasshopper songs are species-specific,
+amplitude-modulated (AM) broad-band acoustic signals 
 
 Among the several thousand recognized grasshopper
 species~(\bcite{cigliano2018orthoptera}), diverse species-specific sound
@@ -148,6 +149,10 @@ Strong dependence on acoustic signals for ranged communication\\
 - Mate attraction/evaluation, rival deterrence, loss-of-signal predator alarm\\
 $\rightarrow$ Elaborate acoustic behaviors co-depend on reliable auditory perception
 
+
+
+
+
 Songs = Amplitude-modulated (AM) broad-band acoustic signals\\
 - Generated by stridulatory movement of hindlegs against forewings\\
 - Shorter time scales: Characteristic temporal waveform pattern\\