diff --git a/cite.bib b/cite.bib
index 5f515d4..1f799ea 100644
--- a/cite.bib
+++ b/cite.bib
@@ -213,6 +213,17 @@
publisher={Springer}
}# Cited
+@article{helversen1998acoustic,
+ title={Acoustic pattern recognition in a grasshopper: processing in the time or frequency domain?},
+ author={von Helversen, Dagmar and von Helversen, Otto},
+ journal={Biological Cybernetics},
+ volume={79},
+ number={6},
+ pages={467--476},
+ year={1998},
+ publisher={Springer}
+}# Cited
+
@article{helversen2004acoustic,
title={{Acoustic communication in a duetting grasshopper: Receiver response variability, male strategies and signal design}},
author={von Helversen, Dagmar and Balakrishnan, Rohini and von Helversen, Otto},
diff --git a/literature/Helversen_Helversen_1998.pdf b/literature/Helversen_Helversen_1998.pdf
new file mode 100644
index 0000000..a6dfca2
Binary files /dev/null and b/literature/Helversen_Helversen_1998.pdf differ
diff --git a/main.aux b/main.aux
index ff53a7b..b4f79a1 100644
--- a/main.aux
+++ b/main.aux
@@ -46,6 +46,10 @@
\abx@aux@segm{0}{0}{stumpner1994song}
\abx@aux@cite{0}{helversen1997recognition}
\abx@aux@segm{0}{0}{helversen1997recognition}
+\abx@aux@cite{0}{helversen1993absolute}
+\abx@aux@segm{0}{0}{helversen1993absolute}
+\abx@aux@cite{0}{helversen1997recognition}
+\abx@aux@segm{0}{0}{helversen1997recognition}
\abx@aux@page{7}{2}
\abx@aux@page{8}{2}
\abx@aux@page{9}{2}
@@ -59,6 +63,8 @@
\abx@aux@page{17}{2}
\abx@aux@page{18}{2}
\abx@aux@page{19}{2}
+\abx@aux@page{20}{2}
+\abx@aux@page{21}{2}
\abx@aux@cite{0}{clemens2013computational}
\abx@aux@segm{0}{0}{clemens2013computational}
\abx@aux@cite{0}{clemens2013feature}
@@ -73,9 +79,9 @@
\abx@aux@segm{0}{0}{rehbein1976auditory}
\abx@aux@cite{0}{eichendorf1980projections}
\abx@aux@segm{0}{0}{eichendorf1980projections}
-\abx@aux@page{20}{3}
-\abx@aux@page{21}{3}
\abx@aux@page{22}{3}
+\abx@aux@page{23}{3}
+\abx@aux@page{24}{3}
\abx@aux@cite{0}{clemens2011efficient}
\abx@aux@segm{0}{0}{clemens2011efficient}
\abx@aux@cite{0}{clemens2011efficient}
@@ -83,13 +89,13 @@
\abx@aux@cite{0}{clemens2011efficient}
\abx@aux@segm{0}{0}{clemens2011efficient}
\@writefile{toc}{\contentsline {section}{\numberline {2}Developing a functional model of\\the grasshopper auditory pathway}{4}{}\protected@file@percent }
-\abx@aux@page{23}{4}
-\abx@aux@page{24}{4}
\abx@aux@page{25}{4}
\abx@aux@page{26}{4}
\abx@aux@page{27}{4}
\abx@aux@page{28}{4}
\abx@aux@page{29}{4}
+\abx@aux@page{30}{4}
+\abx@aux@page{31}{4}
\abx@aux@cite{0}{michelsen1971frequency}
\abx@aux@segm{0}{0}{michelsen1971frequency}
\abx@aux@cite{0}{windmill2008time}
@@ -103,11 +109,11 @@
\@writefile{lof}{\contentsline {figure}{\numberline {1}{\ignorespaces Grasshopper auditory system}}{5}{}\protected@file@percent }
\newlabel{fig:pathway}{{1}{5}{}{}{}}
\@writefile{toc}{\contentsline {subsection}{\numberline {2.1}Population-driven signal pre-processing}{5}{}\protected@file@percent }
-\abx@aux@page{30}{5}
-\abx@aux@page{31}{5}
\abx@aux@page{32}{5}
-\newlabel{eq:bandpass}{{1}{5}{}{}{}}
\abx@aux@page{33}{5}
+\abx@aux@page{34}{5}
+\newlabel{eq:bandpass}{{1}{5}{}{}{}}
+\abx@aux@page{35}{5}
\abx@aux@cite{0}{suga1960peripheral}
\abx@aux@segm{0}{0}{suga1960peripheral}
\abx@aux@cite{0}{gollisch2002energy}
@@ -118,14 +124,14 @@
\abx@aux@segm{0}{0}{clemens2010intensity}
\abx@aux@cite{0}{fisch2012channel}
\abx@aux@segm{0}{0}{fisch2012channel}
-\abx@aux@page{34}{6}
-\newlabel{eq:env}{{2}{6}{}{}{}}
-\abx@aux@page{35}{6}
\abx@aux@page{36}{6}
-\newlabel{eq:log}{{3}{6}{}{}{}}
+\newlabel{eq:env}{{2}{6}{}{}{}}
\abx@aux@page{37}{6}
\abx@aux@page{38}{6}
+\newlabel{eq:log}{{3}{6}{}{}{}}
\abx@aux@page{39}{6}
+\abx@aux@page{40}{6}
+\abx@aux@page{41}{6}
\newlabel{eq:highpass}{{4}{6}{}{}{}}
\@writefile{toc}{\contentsline {subsection}{\numberline {2.2}Feature extraction by individual neurons}{6}{}\protected@file@percent }
\newlabel{eq:gabor}{{5}{6}{}{}{}}
diff --git a/main.bbl-SAVE-ERROR b/main.bbl-SAVE-ERROR
index 3bd99ea..4e4cc1d 100644
--- a/main.bbl-SAVE-ERROR
+++ b/main.bbl-SAVE-ERROR
@@ -19,47 +19,50 @@
\refsection{0}
\datalist[entry]{nyt/global//global/global}
- \entry{cigliano2018orthoptera}{misc}{}
+ \entry{cigliano2024orthoptera}{article}{}
\name{author}{4}{}{%
- {{un=0,uniquepart=base,hash=c6b4d82bb7320d1c96abe22a473cdb96}{%
+ {{un=0,uniquepart=base,hash=55d574c2f3ce70a83e88c1f8180a21ad}{%
family={Cigliano},
familyi={C\bibinitperiod},
- given={MM},
- giveni={M\bibinitperiod},
+ given={María\bibnamedelima Marta},
+ giveni={M\bibinitperiod\bibinitdelim M\bibinitperiod},
givenun=0}}%
- {{un=0,uniquepart=base,hash=7a9031029590d7322b0c5d4e89cfbf83}{%
+ {{un=0,uniquepart=base,hash=ddc53378425a2bcedc93b5b8a7631b14}{%
family={Braun},
familyi={B\bibinitperiod},
- given={H},
+ given={Holger},
giveni={H\bibinitperiod},
givenun=0}}%
- {{un=0,uniquepart=base,hash=aaaccfa9ccf909d7d895cd27982ba973}{%
+ {{un=0,uniquepart=base,hash=4f0a9b90cb2a0e92009f1d443f98ab57}{%
family={Eades},
familyi={E\bibinitperiod},
- given={DC},
- giveni={D\bibinitperiod},
+ given={David\bibnamedelima C},
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givenun=0}}%
- {{un=0,uniquepart=base,hash=88f6127c6ecc7f9498c1c9eeae6131ab}{%
+ {{un=0,uniquepart=base,hash=55914eb1a77ba83ad9377c23563a627c}{%
family={Otte},
familyi={O\bibinitperiod},
- given={D},
+ given={Daniel},
giveni={D\bibinitperiod},
givenun=0}}%
}
- \strng{namehash}{0c8d631f158afb252f6587aa676421b1}
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- \strng{authornamehash}{0c8d631f158afb252f6587aa676421b1}
- \strng{authorfullhash}{ba9abad1da3577c16ef730eadfcd854a}
+ \list{publisher}{1}{%
+ {Species File Group}%
+ }
+ \strng{namehash}{a1bf1dcc4da503ffb332580e758475eb}
+ \strng{fullhash}{a6fbb755de1ae39d9edb48a9a14d6b51}
+ \strng{bibnamehash}{a1bf1dcc4da503ffb332580e758475eb}
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+ \strng{authornamehash}{a1bf1dcc4da503ffb332580e758475eb}
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\field{sortinit}{C}
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\field{labeldatesource}{}
\field{labelnamesource}{author}
\field{labeltitlesource}{title}
- \field{title}{{Orthoptera Species File. Version 5.0/5.0}}
- \field{year}{2018}
+ \field{title}{Orthoptera species file}
+ \field{year}{2024}
\endentry
\entry{clemens2013computational}{article}{}
\name{author}{2}{ul=2}{%
@@ -422,6 +425,7 @@
\strng{authorbibnamehash}{60188e4250a05e56a4e98d5ef5e2452d}
\strng{authornamehash}{60188e4250a05e56a4e98d5ef5e2452d}
\strng{authorfullhash}{60188e4250a05e56a4e98d5ef5e2452d}
+ \field{extraname}{1}
\field{sortinit}{H}
\field{sortinithash}{23a3aa7c24e56cfa16945d55545109b5}
\field{extradatescope}{labelyear}
@@ -436,6 +440,70 @@
\field{pages}{381\bibrangedash 422}
\range{pages}{42}
\endentry
+ \entry{helversen1984parallel}{article}{}
+ \name{author}{1}{}{%
+ {{un=1,uniquepart=given,hash=60188e4250a05e56a4e98d5ef5e2452d}{%
+ family={Helversen},
+ familyi={H\bibinitperiod},
+ given={Dagmar},
+ giveni={D\bibinitperiod},
+ givenun=1,
+ prefix={von},
+ prefixi={v\bibinitperiod},
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+ }
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+ \field{labeldatesource}{}
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+ \field{labeltitlesource}{title}
+ \field{journaltitle}{J Comp Physiol A}
+ \field{title}{{Parallel processing in auditory pattern recognition and directional analysis by the grasshopper Chorthippus biguttulus L. (Acrididae)}}
+ \field{volume}{154}
+ \field{year}{1984}
+ \field{pages}{837\bibrangedash 846}
+ \range{pages}{10}
+ \endentry
+ \entry{helversen1993absolute}{article}{}
+ \name{author}{1}{}{%
+ {{un=1,uniquepart=given,hash=60188e4250a05e56a4e98d5ef5e2452d}{%
+ family={Helversen},
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+ given={Dagmar},
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+ \field{labeltitlesource}{title}
+ \field{journaltitle}{J Comp Physiol A}
+ \field{title}{{‘Absolute steepness’ of ramps as an essential cue for auditory pattern recognition by a grasshopper (Orthoptera; Acrididae; Chorthippus biguttulus L.)}}
+ \field{volume}{172}
+ \field{year}{1993}
+ \field{pages}{633\bibrangedash 639}
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+ \endentry
\entry{helversen1997recognition}{article}{}
\name{author}{2}{ul=2}{%
{{un=1,uniquepart=given,hash=60188e4250a05e56a4e98d5ef5e2452d}{%
@@ -630,6 +698,46 @@
\field{pages}{103\bibrangedash 141}
\range{pages}{39}
\endentry
+ \entry{kohler2017morphological}{article}{}
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+ \field{journaltitle}{PLoS ONE}
+ \field{title}{{Morphological and colour morph clines along an altitudinal gradient in the meadow grasshopper Pseudochorthippus parallelus}}
+ \field{volume}{12}
+ \field{year}{2017}
+ \field{pages}{e0189815}
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\entry{machens2001discrimination}{article}{}
\name{author}{5}{ul=2}{%
{{un=0,uniquepart=base,hash=ffe881e395423136438b1d8d31ae69bc}{%
@@ -919,8 +1027,48 @@
\field{pages}{263\bibrangedash 280}
\range{pages}{18}
\endentry
+ \entry{ronacher1986routes}{article}{}
+ \name{author}{3}{ul=2}{%
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+ \field{labeltitlesource}{title}
+ \field{journaltitle}{J Comp Physiol A}
+ \field{title}{{Routes and stations in the processing of auditory directional information in the CNS of a grasshopper, as revealed by surgical experiments}}
+ \field{volume}{158}
+ \field{year}{1986}
+ \field{pages}{363\bibrangedash 374}
+ \range{pages}{12}
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\entry{ronacher2015computational}{article}{}
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{{un=0,uniquepart=base,hash=4ce9349f95ced7a4831a5efa0fe5b00f}{%
family={Ronacher},
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giveni={J\bibinitperiod},
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@@ -959,6 +1107,37 @@
\field{pages}{61\bibrangedash 71}
\range{pages}{11}
\endentry
+ \entry{rowell1972variable}{incollection}{}
+ \name{author}{1}{}{%
+ {{un=0,uniquepart=base,hash=546111b4ec3ef736f47e37604251cfdb}{%
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+ {Academic Press}%
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+ \field{title}{The variable coloration of the acridoid grasshoppers.}
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+ \field{year}{1972}
+ \field{pages}{145\bibrangedash 198}
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@@ -999,6 +1178,43 @@
\field{pages}{154\bibrangedash 175}
\range{pages}{22}
\endentry
+ \entry{stumpner1994song}{article}{}
+ \name{author}{2}{}{%
+ {{un=0,uniquepart=base,hash=a7bef35fb61ea64e0f04d6c87e5aa622}{%
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+ \field{labeldatesource}{}
+ \field{labelnamesource}{author}
+ \field{labeltitlesource}{title}
+ \field{journaltitle}{Bioacoustics}
+ \field{title}{{Song production and song recognition in a group of sibling grasshopper species (Chorthippus dorsatus, Ch. dichrous and Ch. loratus: Orthoptera, Acrididae)}}
+ \field{volume}{6}
+ \field{year}{1994}
+ \field{pages}{1\bibrangedash 23}
+ \range{pages}{23}
+ \endentry
\entry{suga1960peripheral}{article}{}
\name{author}{1}{}{%
{{un=0,uniquepart=base,hash=cb8492490b7b1d1c021ccd740577eab6}{%
@@ -1027,6 +1243,80 @@
\field{pages}{533\bibrangedash 546}
\range{pages}{14}
\endentry
+ \entry{tarasova2021eurasius}{article}{}
+ \name{author}{3}{}{%
+ {{un=0,uniquepart=base,hash=c300ab24404f1989e04bf78f5abff394}{%
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+ givenun=0}}%
+ {{un=0,uniquepart=base,hash=9fc45c46207f6085c8e82f85145c8810}{%
+ family={Vedenina},
+ familyi={V\bibinitperiod},
+ given={Varvara\bibnamedelima Yu},
+ giveni={V\bibinitperiod\bibinitdelim Y\bibinitperiod},
+ givenun=0}}%
+ }
+ \strng{namehash}{59594c348caf53965237e6751047e29b}
+ \strng{fullhash}{4019db0904a4652861a5e46c60e16441}
+ \strng{bibnamehash}{4019db0904a4652861a5e46c60e16441}
+ \strng{authorbibnamehash}{4019db0904a4652861a5e46c60e16441}
+ \strng{authornamehash}{59594c348caf53965237e6751047e29b}
+ \strng{authorfullhash}{4019db0904a4652861a5e46c60e16441}
+ \field{sortinit}{T}
+ \field{sortinithash}{9af77f0292593c26bde9a56e688eaee9}
+ \field{extradatescope}{labelyear}
+ \field{labeldatesource}{}
+ \field{labelnamesource}{author}
+ \field{labeltitlesource}{title}
+ \field{journaltitle}{Zootaxa}
+ \field{title}{{Songs and morphology in grasshoppers of the Stenobothrus eurasius group (Orthoptera: Acrdidae: Gomphocerinae) from Russia and adjacent countries: clarifying of taxonomic status}}
+ \field{volume}{4965}
+ \field{year}{2021}
+ \field{pages}{244\bibrangedash 260}
+ \range{pages}{17}
+ \endentry
+ \entry{tishechkin2016acoustic}{article}{}
+ \name{author}{2}{}{%
+ {{un=0,uniquepart=base,hash=7a0671a3469a530c34c88e8031f8a0df}{%
+ family={Tishechkin},
+ familyi={T\bibinitperiod},
+ given={D\bibnamedelima Yu},
+ giveni={D\bibinitperiod\bibinitdelim Y\bibinitperiod},
+ givenun=0}}%
+ {{un=0,uniquepart=base,hash=f785a95dae75fb458ea9c6207b0573e0}{%
+ family={Vedenina},
+ familyi={V\bibinitperiod},
+ given={V\bibnamedelima Yu},
+ giveni={V\bibinitperiod\bibinitdelim Y\bibinitperiod},
+ givenun=0}}%
+ }
+ \strng{namehash}{6c452033787fcbb231e7d6fa62192f39}
+ \strng{fullhash}{6c452033787fcbb231e7d6fa62192f39}
+ \strng{bibnamehash}{6c452033787fcbb231e7d6fa62192f39}
+ \strng{authorbibnamehash}{6c452033787fcbb231e7d6fa62192f39}
+ \strng{authornamehash}{6c452033787fcbb231e7d6fa62192f39}
+ \strng{authorfullhash}{6c452033787fcbb231e7d6fa62192f39}
+ \field{sortinit}{T}
+ \field{sortinithash}{9af77f0292593c26bde9a56e688eaee9}
+ \field{extradatescope}{labelyear}
+ \field{labeldatesource}{}
+ \field{labelnamesource}{author}
+ \field{labeltitlesource}{title}
+ \field{journaltitle}{Entomol Rev}
+ \field{title}{Acoustic signals in insects: A reproductive barrier and a taxonomic character}
+ \field{volume}{96}
+ \field{year}{2016}
+ \field{pages}{1127\bibrangedash 1164}
+ \range{pages}{38}
+ \endentry
\entry{vedenina2011speciation}{article}{}
\name{author}{2}{}{%
{{un=0,uniquepart=base,hash=9fc45c46207f6085c8e82f85145c8810}{%
diff --git a/main.bcf b/main.bcf
index b45e726..eec68dc 100644
--- a/main.bcf
+++ b/main.bcf
@@ -2381,26 +2381,28 @@
helversen1977stridulatory
stumpner1994song
helversen1997recognition
- clemens2013computational
- clemens2013feature
- ronacher2015computational
- rehbein1974structure
- kalmring1975afferent
- rehbein1976auditory
- eichendorf1980projections
- clemens2011efficient
- clemens2011efficient
+ helversen1993absolute
+ helversen1997recognition
+ clemens2013computational
+ clemens2013feature
+ ronacher2015computational
+ rehbein1974structure
+ kalmring1975afferent
+ rehbein1976auditory
+ eichendorf1980projections
clemens2011efficient
- michelsen1971frequency
- windmill2008time
- malkin2014energy
- machens2001discrimination
- machens2001representation
- suga1960peripheral
- gollisch2002energy
- hildebrandt2009origin
- clemens2010intensity
- fisch2012channel
+ clemens2011efficient
+ clemens2011efficient
+ michelsen1971frequency
+ windmill2008time
+ malkin2014energy
+ machens2001discrimination
+ machens2001representation
+ suga1960peripheral
+ gollisch2002energy
+ hildebrandt2009origin
+ clemens2010intensity
+ fisch2012channel
diff --git a/main.blg b/main.blg
index 7a54514..3fcbb9c 100644
--- a/main.blg
+++ b/main.blg
@@ -1,48 +1,49 @@
[0] Config.pm:307> INFO - This is Biber 2.19
[0] Config.pm:310> INFO - Logfile is 'main.blg'
-[36] biber:340> INFO - === Fr Jan 9, 2026, 16:45:53
-[43] Biber.pm:419> INFO - Reading 'main.bcf'
-[70] Biber.pm:979> INFO - Found 35 citekeys in bib section 0
-[75] Biber.pm:4419> INFO - Processing section 0
-[80] Biber.pm:4610> INFO - Looking for bibtex file 'cite.bib' for section 0
-[81] bibtex.pm:1713> INFO - LaTeX decoding ...
+[36] biber:340> INFO - === Mo Jan 12, 2026, 17:15:07
+[42] Biber.pm:419> INFO - Reading 'main.bcf'
+[69] Biber.pm:979> INFO - Found 35 citekeys in bib section 0
+[74] Biber.pm:4419> INFO - Processing section 0
+[79] Biber.pm:4610> INFO - Looking for bibtex file 'cite.bib' for section 0
+[80] bibtex.pm:1713> INFO - LaTeX decoding ...
[104] bibtex.pm:1519> INFO - Found BibTeX data source 'cite.bib'
-[218] UCollate.pm:68> INFO - Overriding locale 'en-US' defaults 'variable = shifted' with 'variable = non-ignorable'
-[218] UCollate.pm:68> INFO - Overriding locale 'en-US' defaults 'normalization = NFD' with 'normalization = prenormalized'
-[218] Biber.pm:4239> INFO - Sorting list 'nyt/global//global/global' of type 'entry' with template 'nyt' and locale 'en-US'
-[218] Biber.pm:4245> INFO - No sort tailoring available for locale 'en-US'
-[232] bbl.pm:660> INFO - Writing 'main.bbl' with encoding 'UTF-8'
-[239] bbl.pm:763> INFO - Output to main.bbl
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 34, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 49, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 90, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 127, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 136, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 145, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 156, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 167, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 176, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 185, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 196, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 205, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 216, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 242, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 279, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 298, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 315, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 324, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 343, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 360, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 403, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 424, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 433, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 464, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 502, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 511, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 540, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 549, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 566, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 575, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 600, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_H8G7/347c261ec4135a5723bef5c751f5078f_54850.utf8, line 617, warning: 6 characters of junk seen at toplevel
-[239] Biber.pm:133> INFO - WARNINGS: 32
+[216] UCollate.pm:68> INFO - Overriding locale 'en-US' defaults 'normalization = NFD' with 'normalization = prenormalized'
+[216] UCollate.pm:68> INFO - Overriding locale 'en-US' defaults 'variable = shifted' with 'variable = non-ignorable'
+[216] Biber.pm:4239> INFO - Sorting list 'nyt/global//global/global' of type 'entry' with template 'nyt' and locale 'en-US'
+[216] Biber.pm:4245> INFO - No sort tailoring available for locale 'en-US'
+[231] bbl.pm:660> INFO - Writing 'main.bbl' with encoding 'UTF-8'
+[237] bbl.pm:763> INFO - Output to main.bbl
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 34, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 49, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 90, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 127, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 136, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 145, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 156, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 167, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 176, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 185, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 196, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 205, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 216, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 227, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 253, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 290, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 309, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 326, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 335, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 354, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 371, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 414, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 435, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 444, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 475, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 513, warning: 6 characters of junk seen at toplevel
+[237] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 522, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 551, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 560, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 577, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 586, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 611, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:131> WARN - BibTeX subsystem: /tmp/biber_tmp_RZjz/347c261ec4135a5723bef5c751f5078f_83279.utf8, line 628, warning: 6 characters of junk seen at toplevel
+[238] Biber.pm:133> INFO - WARNINGS: 33
diff --git a/main.fdb_latexmk b/main.fdb_latexmk
index 38cccf7..fe19730 100644
--- a/main.fdb_latexmk
+++ b/main.fdb_latexmk
@@ -1,14 +1,14 @@
# Fdb version 4
-["biber main"] 1767973552.97501 "main.bcf" "main.bbl" "main" 1767973577.05757 0
- "cite.bib" 1767972802.86665 21620 2a322b3b3a6d87b5fabeb885b861b90c ""
- "main.bcf" 1767973577.01484 110388 8b6a35c14e086c0afad39ac12247aa4b "pdflatex"
+["biber main"] 1768234506.98088 "main.bcf" "main.bbl" "main" 1768234806.56873 0
+ "cite.bib" 1768229004.16758 21945 8fee1637f6d42a0b5ea12f6f42cc2ecc ""
+ "main.bcf" 1768234806.52367 110545 53b67a0ba6714cf08c1a6240bb315881 "pdflatex"
(generated)
"main.bbl"
"main.blg"
(rewritten before read)
-["pdflatex"] 1767973576.39041 "/home/hartling/phd/paper/paper_2025/main.tex" "main.pdf" "main" 1767973577.05778 0
+["pdflatex"] 1768234805.88012 "/home/hartling/phd/paper/paper_2025/main.tex" "main.pdf" "main" 1768234806.56895 0
"/etc/texmf/web2c/texmf.cnf" 1761560044.43676 475 c0e671620eb5563b2130f56340a5fde8 ""
- "/home/hartling/phd/paper/paper_2025/main.tex" 1767973576.31685 25860 e708fe3bb04bb676b3eb0719957a874d ""
+ "/home/hartling/phd/paper/paper_2025/main.tex" 1768234805.80468 26319 d509a6a4de3a0f61df6d7bd9954d0a09 ""
"/usr/share/texlive/texmf-dist/fonts/map/fontname/texfonts.map" 1577235249 3524 cb3e574dea2d1052e39280babc910dc8 ""
"/usr/share/texlive/texmf-dist/fonts/tfm/public/amsfonts/cmextra/cmex7.tfm" 1246382020 1004 54797486969f23fa377b128694d548df ""
"/usr/share/texlive/texmf-dist/fonts/tfm/public/amsfonts/cmextra/cmex8.tfm" 1246382020 988 bdf658c3bfc2d96d3c8b02cfc1c94c20 ""
@@ -131,10 +131,10 @@
"/var/lib/texmf/web2c/pdftex/pdflatex.fmt" 1761648508 8213325 7fd20752ab46ff9aa583e4973d7433df ""
"figures/fig_auditory_pathway.pdf" 1764926286.29522 997047 a3424a5b5dfa1dfb4f650bcde9c035bb ""
"figures/fig_auditory_pathway.pdf_tex" 1764926286.29622 2096 aa44b3485835aa1bb41439bee4b9c721 ""
- "main.aux" 1767973577.01284 6273 1d04cd647369114f59aa168b9ce2136b "pdflatex"
- "main.bbl" 1767973553.50005 58057 6e997a5d86e425212218bc8e990ec5b3 "biber main"
- "main.run.xml" 1767973577.01584 2305 d0067550f22ade5b49f22d40cea22838 "pdflatex"
- "main.tex" 1767973576.31685 25860 e708fe3bb04bb676b3eb0719957a874d ""
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+This is pdfTeX, Version 3.141592653-2.6-1.40.25 (TeX Live 2023/Debian) (preloaded format=pdflatex 2025.10.28) 12 JAN 2026 17:20
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@@ -651,29 +651,14 @@ Overfull \hbox (22.7829pt too wide) in paragraph at lines 93--123
[]
-Overfull \hbox (24.27313pt too wide) in paragraph at lines 93--123
-\OT1/cmr/m/n/12 sulted in a mas-sive species di-ver-si-fi-ca-tion ([], []),
- []
-
-
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-\OT1/cmr/m/n/12 with over 6800 rec-og-nized grasshop-per species in the \OT1/cmr/m/it/12 Acri-di-dae \OT1/cmr/m/n/12 fam-ily ([]).
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[]\OT1/cmr/m/n/12 Similar re-sponse/filter prop-er-ties within re-cep-tor/interneuron pop-u-la-tions ([]Clemens, Kutzki,
[]
@@ -684,12 +669,12 @@ File: figures/fig_auditory_pathway.pdf Graphic file (type pdf)
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[]\OT1/cmr/m/n/12 ) and, to a lesser ex-tent, the re-cep-tors them-selves ([])
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[]\OT1/cmr/bx/n/17.28 Two mech-a-nisms driv-ing the emer-gence of intensity-
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@@ -93,9 +93,8 @@ formalization of the underlying structures and mechanisms.
One sensory system about which extensive information has been gathered over the
years is the auditory system of grasshoppers~(\textit{Acrididae}). Grasshoppers
rely on their sense of hearing primarily for intraspecific communication, which
-includes mate attraction and
-evaluation~(\bcite{helversen1972gesang}, \bcite{helversen1993absolute},
-\bcite{helversen1997recognition}), sender
+includes mate attraction and evaluation~(\bcite{helversen1972gesang},
+\bcite{helversen1993absolute}, \bcite{helversen1997recognition}), sender
localization~(\bcite{helversen1988interaural}), courtship display~(SOURCE),
rival deterrence~(\bcite{greenfield1993acoustic}), and loss-of-signal predator
alarm~(SOURCE). In accordance with this rich behavioral repertoire,
@@ -108,32 +107,37 @@ males of the species --- to potential mates within range. These songs are
usually more characteristic of a species than morphological
traits~(\bcite{tishechkin2016acoustic}, \bcite{tarasova2021eurasius}), which
can vary greatly within species~(\bcite{rowell1972variable},
-\bcite{kohler2017morphological}). The reliance on acoustic signals to mediate
-reproduction represents a strong evolutionary driving force, that resulted in a
-massive species diversification~(\bcite{vedenina2011speciation},
+\bcite{kohler2017morphological}). The reliance on songs to mediate reproduction
+represents a strong evolutionary driving force, that resulted in a massive
+species diversification~(\bcite{vedenina2011speciation},
\bcite{sevastianov2023evolution}), with over 6800 recognized grasshopper
species in the \textit{Acrididae} family~(\bcite{cigliano2024orthoptera}). It
is this diversity of species, and the crucial role of acoustic communication in
its emergence, that makes the grasshopper auditory system an intriguing
-candidate for attempting to construct a functional model framework. For
-simplicity, we focus on the pathway responsible for the recognition of
-species-specific calling songs, disregarding other auditory functions such as
-directional hearing~(\bcite{helversen1984parallel}, \bcite{ronacher1986routes},
+candidate for attempting to construct a functional model framework. As a
+necessary reduction, the model we propose here focuses on the pathway
+responsible for the recognition of species-specific calling songs, disregarding
+other auditory functions such as directional
+hearing~(\bcite{helversen1984parallel}, \bcite{ronacher1986routes},
\bcite{helversen1988interaural}).
-The characteristic calling songs are
-produced by stridulation, during which the grasshopper pulls the serrated
-stridulatory file on its hindlegs across a resonating vein on the
+To understand the functional challenges faced by the grasshopper auditory
+system, one has to understand the properties of the songs it is designed to
+recognize. Grasshopper songs are amplitude-modulated broad-band acoustic
+signals. Most songs are produced by stridulation, during which the animal pulls
+the serrated stridulatory file on its hindlegs across a resonating vein on the
forewings~(\bcite{helversen1977stridulatory}, \bcite{stumpner1994song},
-\bcite{helversen1997recognition}).
+\bcite{helversen1997recognition}). Every tooth that strikes the vein generates
+a brief pulse of sound. Multiple pulses make up a syllable; and the alternation
+of syllables and relatively quiet pauses forms a characteristic, through noisy,
+waveform pattern. Species-specific song recognition depends on certain temporal
+song parameters, such as the slope of pulse
+onsets~(\bcite{helversen1993absolute}). The amplitude modulation of the song
+already carries sufficient information to allow for successful
+recognition~(\bcite{helversen1997recognition}). Since grasshoppers are
+ectotherms, the temporal structure of their songs warps with temperature.
-Strong dependence on acoustic signals for ranged communication\\
-- Diverse species-specific sound repertoires and production mechanisms\\
-- Different contexts/ranges: Stridulatory, mandibular, wings, walking sounds\\
-- Mate attraction/evaluation, rival deterrence, loss-of-signal predator alarm\\
-$\rightarrow$ Elaborate acoustic behaviors co-depend on reliable auditory perception
-
Songs = Amplitude-modulated (AM) broad-band acoustic signals\\
- Generated by stridulatory movement of hindlegs against forewings\\
- Shorter time scales: Characteristic temporal waveform pattern\\